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Bolboschoenus fluviatilis (Torrey) Soják [family CYPERACEAE]

Flora of North America, Vol 23,
Flora of North America (FNA)
Reference Sources
Culms 100–200 cm × 5–15 mm. Leaves: sheaths reaching to middle of culm or higher, fronts convex (to concave) and papery at mouth, veins reaching apex, very rarely diverging leaving triangular, veinless, membranous area; widest blade 7–22 mm wide. Inflorescences subumbellate, with all or most spikelets solitary or in clusters of 2–3(–8) on 4–12 rays, rays not exceeding 10 cm; involucral bracts that surpass inflorescences 3–6, widest bract 4–15 mm wide. Spikelets 10–40, ovoid to lanceoloid, 10–25 × 6–10 mm, base cuneate to rounded; scales often loosely imbricate, orange-brown to stramineous, usually obscurely lineolate-spotted, 7–10 × 3–4 mm, membranous and translucent, apex 2-fid 0.5–1 mm deep, awns fairly stout, 2–3 × 0.5 mm wide at base. Flowers: perianth bristles tightly attached to shed achene, pale brown, very stout, equaling achene; anthers yellow, 4 mm; styles 3-fid. Achenes grayish or dark brown, often in patches, obovoid to obpyriform, all nearly equilaterally markedly trigonous to slightly compressed, angles equally rounded, 3.8–5.5 × 2–2.9 mm, apex rounded, beak 0.2–0.8 mm, surface rather dull, exocarp cells usually not evident at 20X; in cross section exocarp much thinner than mesocarp and its cells very small, isodiametric; achene specific gravity greater than water. 2n = 94.

Bolboschoenus [family CYPERACEAE]

Flora of North America, Vol 23,
Flora of North America (FNA)
Reference Sources
Herbs, perennial, cespitose or not, rhizomatous. Culms solitary or not, cormose at base [except eastern Asian Bolboschoenus planiculmis (F. Schmidt) T. V. Egerova], sharply trigonous. Leaves basal and cauline; sheaths tubular; ligules absent; blades flat or V-shaped in cross section, prominently keeled on abaxial surface, distally trigonous. Inflorescences terminal, subumbellate, subcorymbose, or capitate; spikelets to 80, mostly 4–10 mm diam.; involucral bracts surpassing inflorescence 1–5, spreading or proximal erect, leaflike. Spikelets: scales 25 or more, spirally arranged, each subtending flower, abaxially puberulent, often glabrescent, tip notched and awned. Flowers bisexual; perianth of 3–6 bristles, straight or curved, shorter to slightly longer than achene, retrorsely spinulose; stamens 3; styles linear, 2–3-fid, base persistent. Achenes biconvex to trigonous, 2.3–5.5 mm, smooth.

Bolboschoenus novae-angliae (Britton) S. G. Smith [family CYPERACEAE]

Flora of North America, Vol 23,
Flora of North America (FNA)
Reference Sources
Culms to 150 cm × 5–10 mm. Leaves: sheaths reaching beyond or to middle of culm, fronts convex, papery at mouth, veins reaching apex; widest blade 6–13 mm wide. Inflorescences subumbellate, all or most spikelets solitary or in clusters of 2–7 on 3–11 rays, rays not exceeding 8 cm; involucral bracts that surpass inflorescences 2–4, widest bract 2–8 mm wide. Spikelets 10–40, ovoid or lanceoloid, 15–40 × 5–6(–8) mm; scales tightly appressed or in fruit usually loosely imbricate, medium to dark orange-brown, lineolate-spotted at 15X, 7–9 × 3–4 mm, papery, nearly opaque, apex 2-fid 0.5 mm deep, awn fairly stout, 2–3 × (0.3–)0.5 mm at base. Flowers: perianth bristles weakly attached to shed achene or some to all falling separately, medium brown, 1/2 to equaling length of achene; anthers dark yellow to orange-yellow, 3.5–5 mm; styles (2–)3-fid. Achenes dark to medium brown, color variable often on same achene, shape variable, often in same plant, markedly to obscurely trigonous with equally rounded angles to much compressed-trigonous with abaxial angle broadly rounded, or some biconvex in same spikelet, obovoid, 3–4.3 × 2.3–3.1 mm, apex rounded to nearly truncate, beak 0.1–0.5 mm, surface glossy or glossy with dull patches, entirely clearly cellular at 20X or faintly cellular to noncellular at 30X, often in patches; in cross section exocarp 1/3–2/3 of mesocarp thickness and its cells variably enlarged, 1.5–3 times deeper than wide; achene specific gravity greater to less than water.

Bolboschoenus robustus (Pursh) Soják [family CYPERACEAE]

Flora of North America, Vol 23,
Flora of North America (FNA)
Reference Sources
Culms 50–150 cm × 4–8 mm. Leaves: sheaths reaching beyond middle of culm, fronts usually convex, papery at mouth, veins reaching margin, or occasionally membranous with veins diverging proximal to apex leaving triangular, veinless area; widest blade 4–12 mm wide. Inflorescences simply branched with 1 or not more than 1/2 of spikelets solitary or in clusters of 2(–3) on 1–7 rays, rays not exceeding 7 cm; involucral bracts that surpass inflorescence 2–4, widest bract 2–7 mm wide. Spikelets (1–)5–25, broadly ovoid to rarely lanceoloid, 10–30 × 6–10 mm, base usually truncate; scales usually tightly imbricate, medium to dark orange-brown, lineolate-spotted at 15X, usually 6–9 × 3–4 mm, papery and nearly opaque, rarely membranous, apex 2-fid, 1 mm deep, awn very stout, 2–3 × 0.5 mm at base. Flowers: perianth bristles not persistent on shed achene or 1–2 weakly attached to shed achene, dark red-brown, 1/2 achene length; anthers brownish orange, 1.5–2.5 mm; styles (2–)3-fid. Achenes dark to medium brown, obovoid, compressed-trigonous with broadly rounded abaxial angle or sometimes biconvex, 2.7–3.5 × 1.8–2.8 mm, apex truncate to broadly rounded, beak 0.1–0.3 mm, surface glossy, exocarp cells evident at 10–20X; in achene cross section exocarp 2 times thicker than mesocarp and its cells greatly enlarged, 3 times or more deeper than wide; achene specific gravity much less than water.

Bolboschoenus maritimus (Linnaeus) Palla [family CYPERACEAE]

Flora of North America, Vol 23,
Flora of North America (FNA)
Reference Sources
Culms 50–150 cm × (1–)3–8 mm. Leaves: sheaths reaching ± to mid culm, fronts usually membranous at mouth, apex concave to convex, veins diverging proximal to summit leaving triangular, veinless area (often disintegrating); widest blade 2–12 mm wide. Inflorescences with all spikelets sessile, simply branched with not more than 1/2 of spikelets solitary or in clusters of 2–10 on 1–4 rays, rays not exceeding 8 cm; involucral bracts that surpass inflorescences 1–4, widest bract 1–6 mm wide. Spikelets (1–)2–40, ovoid to lanceoloid, 7–40 × (4–)7–10 mm, base usually cuneate to rounded; scales often loosely imbricate, bright orange-brown to stramineous, often obscurely lineolate-spotted, 5–8 × 3–4 mm, membranous, translucent, apex acute, 2-fid 0.5–1 mm deep, awn slender, 1–3 × 0.25 mm at base. Flowers: perianth bristles not persistent on shed achenes or sometimes 1–few weakly persistent, medium brown, to 1/2 achene length; anthers yellow, very rarely orange, 2–4 mm; styles 2–3-fid. Achenes medium to dark brown, rarely yellowish or whitish, obovoid, biconvex, or concave-sided, rarely obtusely compressed trigonous, 2.3–4.1 × 1.9–2.8 mm, apex rounded to nearly truncate, beak 0.1–0.4 mm, surface glossy, exocarp (epidermal) cells usually clearly visible at 10–20X; in achene cross section exocarp 2 times as thick as mesocarp, its cells greatly enlarged, at least 3 times deeper than wide; achene specific gravity much less than water.